modify predator M2: keep M2 separate from M1 and add it to Z on area basis

[Rick-Methot-NOAA]

follow-up from issue #45

a predator is a fleet, so operates in a specific area
natural mortality is a biological trait, so currently is not area-specific.
need to make natural mortality area-specific so that each predator can affect M2 in its area.
This means that pred_M2 should not be added to M1 in the biology step.
Instead, pred_M2(f) should be included when Z(p) is calculated within each area.

[k-doering-NOAA]

@Rick-Methot-NOAA , it looks like this is the last outstanding issue according to the projects spreadsheet; will this be worked before 3.30.19? Or do we want to delay it?

[Rick-Methot-NOAA]

attached doc describes the revised logic that allows natM to be area and time specific, which is needed for predators and is needed for future development of transitional SPR (#88 ) calculations.
natM tech doc.txt