LaTeX templates for manuscripts (switchable bioRxiv preprint and journal submission templates), for use on Overleaf.
An ideal manuscript LaTeX template should be able to generate two versions of the manuscript:
- a preprint version - typeset to look like a finished paper
- a journal submission version - better for reviewing, with line numbers and different spacing, etc.
The ideal template should be able to switch between these two manuscript layouts. A preview of both outputs is shown below.
To use, just clone this repo and import into Overleaf or a directory on your own computer to use a local LaTeX installation.
The following files and folders are not required: img, examples, README.md, LICENSE.
Write your manuscript in 01_Article_MainText.tex and then comment either line 2 or 3 of 00_Article_Merge.tex to select between outputs.
There are optional Supplementary tex files that can also be edited.
If they are not required, comment the appropriate lines in 00_Article_Merge.tex.
Tested on Overleaf - TeXLive 2025 with pdfLaTeX compiler.
The initial overleaf template was forked from the template created by Henriques lab available here. This is a modification of the PNAS template (also available on Overleaf) which is widely used on bioRxiv. We have made a few changes to the HenriquesLab bioRxiv template and made it possible to easily generate a journal submission version.
Changes include:
- genuine bioRxiv logo used in the footer
- support for ORCiDs using
\orcidlink - Helvetica for readability
- greater range of custom units
- simplified directory structure
The provided files (with comments) provide guidance on how to put your manuscript together. There are a few things to be aware of:
- the manuscript is compiled from separate
.texfiles which are listed in00_Article_Merge.tex. The first included.texfile is the main text (01_Article_MainText.tex) and this has its own reference section, which means that a subsequent "Supplementary Information" section will have a separate reference section. - switching from the preprint version (
\documentclass[twocolumn]{bioRxiv}) to the plain manuscript version (\documentclass[submit]{bioRxiv}) works well, but has some foibles. Any figures that usefigure{}environment rather thanfigure*{}may need resizing by a factor of 0.5 to remain at the intended size. - figures should be prepared as a single file and placed in the
figuresfolder - use of figures compiled from multiple files in LaTeX is not supported.

